The Dark Prince was once a human that was the heir and king of the Castle of Light. For unexplained reasons, he was hated and feared by his subjects. The creatures of Mavolia took advantage of this and made him an offer. In exchange for destroying his own kingdom, he would become their leader. He accepted the offer, and within the Mana Tree died at the hands of his brother while attempting to release the seal on the Mana Gate and fully open the door to Mavolia.